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Alpheus Benthos is the primary antagonist of season 2.
He commands a nuclear-powered submarine of his own design, which he clams to be the fastest sub around. The sub is controlled by an A.I. named A.R.I.A. (Augmented Response Intelligent Application).
Alpheus is the self-programed "nemesis" of Antaeus Nekton, who believes their fates are intertwine. He seeks the secrets of Lemuria, so he can awaken a mysterious doomsday weapon called "The Terror", and is working along-side Proteus to do so.
Alpheus's suit is embedded with electronics that allow him to shoot disabling rays right from his forearms.